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CHAPTER 19 Nutrition/Digestive System
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one Consider, in this case, the large globules of partially digested fat that do not mix very well with the soupy chyme entering the duodenum from the stomach Bile from the liver acts to emulsify the large fat globules, breaking them apart (or, in a sense, milking them out of the rest of the chyme) As a result, a separate foam of tiny fat droplets is created within the small intestine [Study suggestion: Pour some liquid detergent onto a bunch of greasy plates, and then observe what happens In what way does this liquid detergent act somewhat like bile ] Bile is secreted continuously, day and night, into the right and left hepatic (heh-PAT-ik) or liver ducts These ducts carry the bile into the cystic (SIStic) duct Cyst means bladder or sac, while chole (KOH-lee) is Latin for bile or gall Hence, the compound word, cholecyst (KOH-luh-sist), translates into English as gall bladder or bile sac The cholecyst (gall bladder) is a muscular-walled sac that receives the bile from the liver and stores it temporarily When the duodenum becomes swollen with fatty chyme, a hormone is released that stimulates the walls of the gall bladder to contract A load of bile is squirted out of the cholecyst, much like a slug of brownish-green pea soup or gravy being squeezed out of a rubber balloon The bile squirts into the cystic duct, and then into the common bile duct, which carries it the rest of the way down into the duodenum Here, then, the emulsi cation of fat takes place As shown in Figure 194, the pancreatic (pan-kree-AT-ik) duct extends from the pancreas and merges with the base of the common bile duct The pancreatic duct is the main passageway for the pancreatic juice Surrounding both of them at their point of union is the hepatopancreatic (heh-PAT-ohpan-kree-AT-ik) sphincter (SFINK-ter) The hepatopancreatic sphincter is a ring of smooth muscle that regulates the emptying of both the common bile duct (the hepatic or liver portion) and the pancreatic duct into the small intestine When this sphincter (muscular ring) relaxes, bile and pancreatic juice ow through the duodenal (dew-AH-deh-nal) papilla (pah-PIL-lah) The duodenal papilla is a little nipple or pimple (papill) -like projection with a hole in its center Bile from the liver and pancreatic juice from the exocrine gland portion of the pancreas ( 15) drip into the duodenum through the hole in the duodenal papilla The pancreatic juice contains sodium bicarbonate (buy-CAR-buh-nayt), symbolized chemically as NaHCO3, as well as a variety of digestive enzymes These enzymes include amylases (starch-splitters), lipases (LIE-pay-sez) or fat-splitters, and proteases (PROH-tee-ay-sez) or protein-splitters The lipases, for example, complete the chemical digestion of fat or lipids, after they have been emulsi ed by bile into a fatty foam The resulting products,
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PART 4 Anatomy and Physiology of Animals
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LIVER Bile R hepatic duct Cystic duct Cholecyst (gallbladder) Common bile duct Bile L hepatic duct
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STOMACH PANCREATIC DUCT
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Pancreas Common bile duct
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Microvilli VILLUS
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Duodenal papilla Pancreatic duct Presence of twelve (fingerbreadths) Hepatopancreatic sphincter
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Fig 194 The duodenum and its friendly neighbors
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such as fatty acids and the substance, glycerol (GLIH-sir-ahl), are then absorbed across the walls of the small intestine, and into the bloodstream Similarly, the proteases continue the chemical breakdown of proteins into amino acids, which are also absorbed into the bloodstream And the amylases in the small intestine generally nish the chemical breakdown of carbohydrates into simple sugars such as glucose, which are then absorbed Re ecting its critical role in the absorption of nutrients, there are several important modi cations to the mucosa (mew-KOH-sah), the mucous (mucos) membrane lining the small intestine The mucosa of the duodenum, jejunum, and ileum is thrown into thousands of villi (VIL-ee) Review of Figure 194 shows that each single villus (VIL-us) is literally named for its resemblance to a little bump or curved tuft of hair (vill) The surface of each villus, in turn, is covered with dozens of microvilli (MY-kroh-vil-ee) The microvilli are little bumps upon each villus, like tiny tufts of hair The numerous villi and microvilli, by throwing the mucosa up into hundreds of tiny bumps, vastly increases the amount of surface area available for absorption As a result, the absorption of nutrients into the bloodstream from the small intestine is extremely e cient
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