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PART 4 Anatomy and Physiology of Animals
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Renal cortex ( kidney bark ) Renal capsule Renal pyramid
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Renal medulla ( kidney middle )
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Aztec pyramid Renal calyx ( kidney flower cup )
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Ureter RENAL PELVIS ( kidney bowl ) CLOSE-UP OF RENAL CORTEX Glomerulus ( ball of yarn )
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Urine droplets Renal capillaries Urinary tubules
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Fig 201 An overview of renal anatomy
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blunt pyramids constructed by the Aztecs or Inca Indians The tip of each renal pyramid drips urine into a renal calyx (KAY-licks), or kidney ower cup And the urine from each calyx eventually ows into the body of the renal pelvis, before it leaves the kidney via the ureter
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THE URINARY PATHWAY
Figure 202 shows the rest of the urinary pathway, lying beyond the kidney The right and left ureters both dump urine into the urocyst (YUR-oh-sist) or urinary bladder (cyst) The urocyst (urinary bladder) is a hollow, muscularwalled pouch that temporarily stores the urine before it is excreted
CHAPTER 20 Urine and Sex in Animals
UROCYST (URINARY BLADDER) R Ureter L Ureter
Openings for ureters
URINARY SPHINCTER
URETHRA
URINARY ORIFICE (in female)
Fig 202 The urinary pathway
The urocyst empties into the urethra (you-REETH-rah), the tube that helps a person literally make water (urethr) that is, urinate (YUR-ihnayt) Surrounding the upper neck of the urethra is the urinary sphincter Much like the external anal sphincter in the digestive pathway ( 19), the urinary sphincter is a ring of voluntary striated muscle This means, of course, that the contraction and relaxation of this sphincter is under our voluntary control Thus, after we have been adequately potty-trained during early childhood, we can voluntarily relax the urinary sphincter whenever the place and time are right for urination! Finally, urine exits out of the body through the urinary ori ce (OR-ih- s), a tiny, mouth (or)-like opening
The Process of ( Making Water ) or Excreting Urine
We have, from time to time, been using some very basic descriptive equations in this book Let us do so, again We can state the urinary excretion equation, for example:
PART 4 Anatomy and Physiology of Animals
E F R S (This equation is put into visual form within Figure 203) In this equation, E stands for excretion, F for amount ltered, R for the amount of tubular reabsorption, and S for tubular secretion The urinary ltrate (FIL-trayt), or ltration product, comes from the pushing force of the blood pressure against the walls of the renal capillaries in the glomeruli (glah-MEHR-you-lie) This quantity of ltrate is huge, averaging about 180 liters of uid per day, in an average adult! [Study suggestion: Assume that an adult has a blood volume of about 6 liters Then, on average, how many times is this person s entire blood volume ltered out of his glomeruli, each day ] While it may seem wasteful, the huge volume of urinary ltrate (F) acts as the starting point for the urine Because there is so much of this ltrate, the body can adjust many factors to in uence how much urine is actually excreted, under particular current conditions After urinary ltration, one of the chief processes is tubular reabsorption (R) Reabsorption is the movement of material out of the ltrate, across the walls of the urinary tubules, and back into the bloodstream Consider, for example, the tubular reabsorption of glucose Under normal conditions, almost 100% of the glucose that is ltered into the urinary tubules is eventually reabsorbed back into the bloodstream As a result, the urine excreted from the body is nearly free of glucose Several hormones also control the amount of sodium (Na ions) and water (H2O) molecules that are reabsorbed back into the bloodstream Because the amount of sodium (salt) and water reabsorbed can vary greatly, the kidneys play a critical role in regulating the salt water balance of the human bloodstream Under typical conditions, about 99%, or 179 liters, of the urinary ltrate (mostly water) is reabsorbed [Study suggestion: If a person becomes extremely dehydrated, as after excessive sweating, then what do you predict will happen to the amount of H2O reabsorbed Will the percent (%) reabsorbed increase above typical conditions, or decrease below it Why ] Another process, tubular secretion (S), involves the active (ATP-requiring) addition of small quantities of particular chemicals from the bloodstream, into the urinary tubules Molecules of penicillin (pen-ih-SILL-in) and many other antibiotics (an-tih-buy-AH-ticks), for instance, are just too large to be ltered across the walls of the glomeruli Hence, the epithelial cells lining the blood vessels actively pump the penicillin into the urinary tubules Therefore, penicillin is excreted (E) out of the body, via the urine Although only a few milliliters (ml) of uid are generally secreted each day, they still have an important in uence
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