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R Cerebrum L
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Upper motor neuron Central sulcus Directing voluntary movement (goes downward)
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Lower motor neuron Moves the muscul ( mouse ) Body midline
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Fig 1410 Higher motor neurons and the hidden puppeteer
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PART 4 Anatomy and Physiology of Animals
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mass The upper motor neurons situated within the cerebral cortex of the precentral gyrus often direct voluntary movements of the body to occur The precentral gyrus is sometimes called the primary motor area It is here that upper motor neurons decide to initiate the movements of various skeletal muscles (usually on the opposite side of the body) From the primary motor area (precentral gyrus), motor nerve bers descend, cross over the body midline, and then stimulate skeletal muscles on the opposite side of the body to contract To move your right hand, for instance, upper motor neurons within the left precentral gyrus would send down a message, which would eventually cross to the right side of the spinal cord, then out to the muscles in the right hand [Study suggestion: Imagine a mischievous little puppeteer hiding within your left precentral gyrus The strings manipulating his puppet hang down (descend) for some distance These strings are actually nerve bers in a motor pathway After the strings are pulled (excited by a muscle action potential), they activate the puppet (skeletal muscle) to contract]
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Just as motor pathways descend or come down from both higher and lower motor neurons, sensory pathways ascend or rise up from sensory neurons A lower or rst-order sensory neuron is a nerve cell that has part of its dendrites modi ed to form sensory receptors Sensory receptors are specialized nerve endings that are sensitive to a particular kind of goad (poking prod) called a stimulus (STIM-yuh-les) Consider, for example, certain onion-shaped sensory receptors in the skin of the right hand (Figure 1411) They are sensitive to touch or pressure sensations A nerve impulse is created, and then it travels over a peripheral nerve, then a spinal nerve, and into the spinal cord A higher or second-order sensory neuron is then excited Its axon crosses the body midline to the left side of the spinal cord, and then ascends towards the thalamus (THAL-uhmus) The thalamus is an egg-shaped bedroom (thalam) an oval region nestled deep within the cerebrum The thalamus is often described as a sensory relay center This is because sensory nerve impulses (action potentials) are often switched over (or relayed) up onto third-order sensory neurons, whose cell bodies lie within the thalamus And from the thalamus, the axons of the third-order sensory neurons nally carry the information about the stimulus all the way up to the postcentral gyrus
CHAPTER 14 Neuromuscular Connection
As its name clearly indicates, the postcentral gyrus is a raised ring or fold of cerebral cortex that runs up and down just behind or after (post-) the central sulcus The postcentral gyrus is alternately called the primary or general sensory cortex This name re ects the fact that the postcentral gyrus area is the nal destination for most of our general or primary body senses, such as touch, pressure, temperature, vibration, and pain Remember that we imagined there was a hidden puppeteer within the left precentral gyrus, directing much of the motor pathway descending on the right side of the body Likewise, we can draw a sensory homunculus (hohMUNG-kyuh-lus) a little feeling man or dwarf (homuncul ) just above the left postcentral gyrus This little feeling dwarf is a rather bizarre visual method by which anatomists have commonly mapped the nal destinations of sensory nerve bers ascending from various parts on the right side of the body
Sensory homunculus ( little feeling dwarf ) Postcentral gyrus (primary sensory cortex)
Third-order sensory neuron
THALAMUS HYPOTHALAMUS
Cerebellum Higher (second-order) sensory neuron Sends sensory information upward Body midline Stimulus Lower (first-order) sensory neuron
Sensory receptors
Fig 1411 The primary sensory pathway and its little man or dwarf
PART 4 Anatomy and Physiology of Animals
The Cerebellum and Hypothalamus
The thalamus and postcentral gyrus (primary or general sensory cortex) are not the only places in the human brain that receive sensory information Two other very important sensory areas are the cerebellum (sair-uh-BELL-um) or little cerebrum (cerebell) and the hypothalamus (high-poh-THAL-uh-mus) or region below (hypo-) the thalamus (Review Figure 1411) The cerebellum is involved in proprioception (proh-pree-oh-SEP-shun) This long term literally means the process of receiving (-ception) sensations from one s own (propri) self In more direct language, proprioception involves the receiving of sensory stimuli that indicate the relative position of the entire body and its parts within space The sensory receptors for proprioception are often found within the skeletal muscles and joints Thus, you can sense or feel that you are standing upright, or that you have raised your arm up o a table, without even having to look! By providing proprioception, the cerebellum helps the body maintain its vertical balance and reduces the amount of shaking during automatic re ex movements
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