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Integrating Metabolic Pathways ATP Glucose Storage Molecules Metabolic States and Signals Insulin Glucagon Epinephrine Secondary Signals Generalities of Metabolism Phosphorylation Glycogen Metabolic Movements of Glycogen Fat Metabolic Movements of Fat Protein Metabolic Movements of Protein Tissue Cooperation Liver Muscle Adipose Brain Connection of Storage Pools Feeding Fasting Starvation Excitement Interorgan Cycles Cori Cycle Alanine Cycle Ketone Bodies
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Integration of Energy Metabolism
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Welcome to energy metabolism It s the nuts and bolts of life at the cellular level1 Energy metabolism is responsible for maintaining a constant supply of ATP in all different tissues Since some tissues, such as red cells and brain, rely heavily on glucose to make ATP, maintaining the ATP levels in all tissues requires maintaining the availability of glucose The purpose of all the pathways we ll discuss is to maintain ATP and glucose supplies As you prowl around the individual metabolic pathways of energy metabolism, you need to get a feel for the kinds of things you should be looking for as you examine each metabolic scheme There are four individual pathways involved in energy metabolism: glycolysis-gluconeogenesis, fatty acid synthesis oxidation, glycogen synthesis-degradation, and the TCA cycle First look for the overall function of each pathway: What does it do This involves knowing the molecules coming into the pathway and those going out Next, understand when the pathway should be on or off what metabolic states require the pathway to function Then, figure out which tissues use the pathway and how Finally, see how the behavior of the pathway may be integrated into the cooperation between organs and tissues Energy metabolism makes sense if you realize that each individual pathway and each organ has a function Understanding metabolism in every detail may well be impossible, but understanding the general themes is not only possible but important Energy metabolism maintains the supply of ATP and glucose by making storage molecules (glycogen, fat, protein) when food is available and by retrieving ATP and glucose from storage when they are needed The need for glucose or ATP may constitute a demand for massive amounts of immediate energy or it may simply be the need to maintain energy and glucose levels between meals Energy metabolism is regulated in a manner that involves extensive cooperation between different organs, but the goal is always the same: to maintain adequate ATP and glucose levels
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Metabolic pathways interconnect glycogen, fat, and protein reserves to store and retrieve ATP and glucose Most of energy metabolism should make sense don t forget that The function of energy metabolism is reasonably simple all of it is con1
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At least the nuts part is right
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Basic Concepts in Biochemistry
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cerned with putting energy into storage when it s not needed and taking energy out of storage when it is needed It s the details that complicate the picture There are only three types of storage molecules: glycogen, fat, and protein These three storage forms are all connected, and that s what energy metabolism is all about the connections When metabolism works normally, we re assured of a relatively constant supply of energy and glucose No one tissue can survive metabolically without the others Each of the four major tissue types (liver, muscle, adipose, brain) has a specialized metabolic function There are some differences in the metabolic pathways in each tissue; however, these differences are relatively simple and serve to specialize the metabolic functions of the different tissue types There is real cooperation between the different organs Each organ has its own metabolic profile, its own needs, and its own capabilities
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ATP is the immediate source of energy for cellular processes Ultimately, the energy source for all cellular processes is the hydrolysis of ATP There are only two major sources of ATP the TCA cycleelectron transport and glycolysis The major source of ATP is the mitochondrial electron transport chain fueled by the TCA cycle Since mitochondrial electron transport requires oxygen, much of our ATP production is directly linked to the supply of oxygen Red cells, since they don t have any mitochondria, must rely totally on anaerobic glycolysis for energy, and muscle may be forced to rely on anaerobic glycolysis when strenuous exercise uses up oxygen faster than it can be delivered to the muscle When there is no oxygen glycolysis is the only way to make ATP
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