Urine formation in .NET

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Urine formation
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Figure 132 displays the basic mechanism of urine formation that starts with the glomerulus The relatively high BP (blood pressure) within the glomerulus
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Urogenital System
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Fig 132 The basic processes in urine formation and excretion
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creates a powerful pushing force This pushing force results in urinary (YUR-ihnair-ee) or glomerular (gluh-MAHR-you-lar) ltration, abbreviated as F The uid that is ltered out of the bloodstream in the glomerulus passes through the very thin walls of the hollow renal tubules, which run nearby The product is called the urinary ltrate (FIL-trayt) one that (-ate) is ltered ( ltr) A huge volume of urinary ltrate about 180 liters in an average-sized adult is produced every day Obviously, since there are only about 5 to 6 liters of total blood volume, most (about 99%) of the urinary ltrate has to be taken back into the bloodstream [Visualization suggestion: Pretend that you are eating with friends in a restaurant, and you suddenly get the urge to urinate (YUR-ih-nayt) Writing a short note on a napkin, you excuse yourself and go to the restroom Now, what do you predict will happen if you urinate all of the uid present in your urinary (glomerular) ltrate at that time Would you even be able to come back to the table Why could we speculate that the note you wrote was your nal will and testament ] The way we take back 99% of the urinary ltrate is by the process of tubular reabsorption (ree-ab-SORP-shun) Tubular reabsorption is the absorption of
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Life in Land Down Under
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particular chemicals from the urinary ltrate back into the bloodstream again (re-) It is called reabsorption because absorption ( 12) is the way in which chemicals entered the bloodstream from the lumen of the digestive tube in the rst place Consider, for example, the glucose molecule Remember from 12 that glucose molecules are absorbed into the bloodstream after thorough digestion of a ham sandwich Eventually, these glucose molecules leave the bloodstream, enter the urinary ltrate, and are actively reabsorbed (ree-abSORBED) back into the bloodstream again It is almost as if a Magic Chemical Hand dips down into the urinary ltrate The hand uses ATP energy ( 6) to actively scoop up all of the glucose molecules The Hand dumps all of the glucose molecules back into the bloodstream before they have a chance to go down the steep waterfall (be excreted from the body in the urine) Figure 133 depicts this Magic Chemical Hand, which bears the symbolic mark of R, for tubular reabsorption
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R ATP energy used
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Bloodstream Active glucose reabsorption
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Glucose molecule
ry ina Ur rate filt
Excreted urine (contains no glucose)
Fig 133 The magic hand of glucose reabsorption (R)
Urogenital System
Of course, most of the urinary ltrate is water, so a huge number of H2 O molecules (along with various other particles, like Na+ and Cl ions) are also reabsorbed back into the bloodstream Another imaginary hand is also in operation This is the Magic Mini-Hand of tubular secretion (S) Tubular secretion is the process of actively secreting tiny (mini-) amounts of certain substances out across the walls of the blood vessels and into the kidney tubules It is a way of helping the body excrete small quantities of particles that may be present in excessive amounts (such as H+ ions), as well as large antibiotic drug molecules (such as penicillin) that are simply too large to be ltered Figure 134 provides a metaphor of the Magic
Bloodstream
ATP energy used
Active tubular secretion of H+ ions from bloodstream into urinary filtrate
Urinary filtrate
Acid excreted in urine
Fig 134 The magic mini-hand of tubular secretion of H+ ions
PART 6
Life in Land Down Under
Mini-Hand in operation You can see that it is actively scooping individual H+ ions out of the bloodstream, then dumping them into the urinary ltrate Eventually, the H+ ions go down the imaginary waterfall (get excreted from the body within the urine) Even though only a few milliliters of uid are normally secreted out of the bloodstream each day, this is still important in helping prevent acidosis and other metabolic imbalances After the three renal processes of glomerular (urinary) ltration (F), tubular reabsorption (R), and tubular secretion (S) have occurred, the overall result is urine excretion (E) Summarizing the preceding block of information yields the urinary excretion equation:
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