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12. Diagram the results that Meselson and Stahl would have obtained (a) if DNA replication were conservative and (b) if it were dispersive. 13. What type of photo would J. Cairns have obtained if DNA replication were conservative Dispersive
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*Answers to selected exercises and problems are on page A-10.
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Tamarin: Principles of Genetics, Seventh Edition
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III. Molecular Genetics
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9. Chemistry of the Gene1
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The McGraw Hill Companies, 2001
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14. Following is a section of a single strand of DNA. Supply a strand, by the rules of complementarity, that would turn this into a double helix. What RNA bases would primase use if this segment initiated an Okazaki fragment In which direction would replication proceed 5 -ATTCTTGGCATTCGC-3 15. What is a primosome in E. coli a replisome What enzymes make up each What is the relationship between these structures 16. What are the differences between continuous and discontinuous DNA replication Why do both exist 17. Describe the synthesis of an Okazaki fragment. 18. Describe the enzymology of the origin, continuation, and termination of DNA replication in E. coli. 19. Can you think of any other mechanisms besides topoisomerase activity that could release supercoiling in replicating DNA 20. Draw a diagram showing how topoisomerase II (gyrase) might work. 21. Retroviruses are single-stranded RNA viruses that insert their genomes into the host DNA during their life cycle. But only double-stranded DNA can be inserted into double-stranded DNA.
a. Propose a mechanism that retroviruses could use to insert their genomes. b. What novel enzymes might such viruses require 22. Propose a mechanism by which a single strand of DNA can make multiple copies of itself. 23. Progeria is a human disorder that causes affected individuals to age prematurely; a nine-year-old often resembles a sixty- to seventy-year-old individual in appearance and physiology. Suppose you extract DNA from a progeric patient and nd mostly small DNA fragments rather than the expected long DNA molecules. What enzyme(s) might be defective in patients with progeria
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24. Under what circumstances would you expect to see a DNA theta structure D-loop rolling-circle bubbles What function does each structure serve
EUKARYOTIC DNA REPLICATION
25. In developing sea urchins, just after fertilization, the cells divide every thirty to forty minutes. In the adult, the cells divide once every ten to fteen hours. The amount of DNA per cell is the same in each case, but the DNA obviously replicates much faster in developing cells. Propose an explanation to account for the difference in replication time.
C R I T I C A L
T H I N K I N G
Q U E S T I O N S
1. Mutants are used to study various aspects of the phenotype and genotype. How can we study genes that are critically important in the functioning of an organism For example, how do we study mutations in the gene for DNA polymerase III in E. coli, when changes in this gene are usually lethal Remember, to study the genes in bacteria, we need the bacteria to grow and form colonies in order to be scored for their phenotypes.
2. DNA and RNA differ in two major ways: DNA has deoxyribose sugar, whereas RNA has ribose, and DNA has thymine, whereas RNA has uracil. Why might those differences exist other than accidents of evolution
Suggested Readings for chapter 9 are on page B-5.
Tamarin: Principles of Genetics, Seventh Edition
III. Molecular Genetics
10. Gene Expression: Transcription
The McGraw Hill Companies, 2001
GENE EXPRESSION
Transcription
STUDY OBJECTIVES
1. To examine the types of RNA and their roles in gene expression 245, 256 2. To look at the process of transcription, including start and stop signals, in both prokaryotes and eukaryotes 246 3. To investigate posttranscriptional changes in eukaryotic messenger RNAs, including an analysis of intron removal 260
STUDY OUTLINE
Types of RNA 245 Prokaryotic DNA Transcription 246 DNA-RNA Complementarity 246 Prokaryotic RNA Polymerase 247 Prokaryotic Initiation and Termination Signals for Transcription 248 Ribosomes and Ribosomal RNA 256 Transfer RNA 256 Similarities of All Transfer RNAs 257 Transfer RNA Loops 258 Eukaryotic DNA Transcription 260 The Nucleolus in Eukaryotes 260 Differences Between Eukaryotic and Prokaryotic Transcription 261 Promoters 262 Caps and Tails 265 Introns 265 RNA Editing 275 Updated Information About the Flow of Genetic Information 275 Reverse Transcription 276 RNA Self-Replication 276 DNA Involvement in Translation 276 Summary 277 Solved Problems 277 Exercises and Problems 278 Critical Thinking Questions 279 Box 10.1 Observing Transcription in Real Time 250 Box 10.2 Polymerase Collisions: What Can a Cell Do 252 Box 10.3 Are Viroids Escaped Introns 272
A computer model of the serine transfer RNA. The amino acid binding site is yellow; the anticodon is red. ( Ken Eward/SPL/Photo Researchers.)
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