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12. The histidine operon is a repressible operon. The corepressor is charged tRNAHis, and its gene is not part of the operon. For the following mutants, tell whether the enzymes of the operon will be made; then tell whether each mutant would be cis-dominant in a partial diploid. a. RNA polymerase cannot bind the promoter. b. The repressor-corepressor complex cannot bind operator DNA (the operator has the normal sequence). c. The repressor cannot bind charged tRNAHis.
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22. Why are IS elements sometimes referred to as sel sh DNA 23. What are the differences among an IS element, a transposon, an intron, a plasmid, and a cointegrate 24. Describe the Shapiro model of transposition. What are the roles of transposase, DNA polymerase I, ligase, and resolvase 25. Why are transposons anked by direct repeats 26. How do transposons induce deletions inversions 27. Describe how a transposon controls the expression of the agellar phase in Salmonella. 28. What is a polar mutation What can cause it
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13. Describe the interaction of the attenuator and the operator control mechanisms in the trp operon of E. coli under varying concentrations of tryptophan in the cell. How does attenuator control react to shortages of other amino acids
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29. List the steps from transcription through translation to enzyme function, noting all the points at which control could be exerted. (See also TRANSLATIONAL CONTROL and POSTTRANSLATIONAL CONTROL)
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Tamarin: Principles of Genetics, Seventh Edition
III. Molecular Genetics
14. Gene Expression: Control in Prokaryotes and Phages
The McGraw Hill Companies, 2001
Fourteen
Gene Expression: Control in Prokaryotes and Phages
30. What are the advantages of transcriptional control over translational control (See also TRANSLATIONAL CONTROL) 31. How are heat shock proteins induced 32. In phage T4, the genes rIIA and rIIB lie adjacent to each other on the T4 chromosome. During the early phase of infection, rIIA and rIIB products are present in equimolar amounts. In the late phase of infection, the amount of rIIB protein is ten to fteen times higher than that of rIIA protein. Nonsense mutations (mutations to a stop codon) in rIIA eliminate early but not late rIIB transcription. In the mutants that contain small deletions near the end of rIIA, the amount of rIIA product is always equal to the amount of rIIB product, regardless of the time of infection. Based on this information, devise a map of the rII region. Include the location(s) of the promoter(s).
TRANSLATIONAL CONTROL
33. What is the stringent response How does it work What is an idling reaction 34. What is antisense RNA How does it work What is the obvious source of this regulatory RNA How could this RNA be used to treat a disease clinically
POSTTRANSLATIONAL CONTROL
35. What is feedback inhibition What other roles do allosteric proteins play in regulating gene expression 36. What controls the rate of degradation of proteins
C R I T I C A L
T H I N K I N G
Q U E S T I O N S
1. From an evolutionary perspective, why do you think Escherichia coli evolved a CAP system of positive control of gene expression Why not just metabolize any and all sugars in the environment as they appear
2. Why might some proteins and messenger RNAs produced in Escherichia coli be degraded so quickly
Suggested Readings for chapter 14 are on page B-13.
Tamarin: Principles of Genetics, Seventh Edition
III. Molecular Genetics
15. The Eukaryotic Chromosome
The McGraw Hill Companies, 2001
THE EUKARYOTIC CHROMOSOME
STUDY OBJECTIVES
1. To examine the arrangement of DNA and proteins comprising the eukaryotic chromosome 440 2. To look at the nature of centromeres and telomeres in eukaryotic chromosomes 453 3. To analyze the nature of the DNA in eukaryotic chromosomes 457
STUDY OUTLINE
The Eukaryotic Cell 440 The Eukaryotic Chromosome 440 DNA Arrangement 440 Nucleoprotein Composition 442 Chromosomal Banding 451 Centromeres and Telomeres 453 The C-Value Paradox 457 Summary 461 Solved Problems 462 Exercises and Problems 462 Critical Thinking Questions 463 Box 15.1 How Big Is Big, How Small Is Small 441 Box 15.2 High-Speed Chromosomal Sorting 444
Arti cially colored scanning electron micrograph of part of a polytene salivary gland chromosome from a fruit y (Drosophila), revealing the underlying banding pattern. ( Professors P. Motta and T. Naguro/
SPL/Photo Researchers. inc.)
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