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The McGraw Hill Companies, 2001
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incomplete dominance The situation in which both alleles of the heterozygote in uence the phenotype. The phenotype is usually intermediate between the two homozygous forms. independent assortment, rule of Mendel s second rule, describing the independent segregation of alleles of different loci. inducible system A system (a coordinated group of enzymes involved in a catabolic pathway) is inducible if the metabolite it works upon causes transcription of the genes controlling these enzymes. These systems are primarily prokaryotic operons. induction In regard to temperate phages, the process of causing a prophage to become virulent. informatics See bioinformatics. initiation codon The messenger RNA sequence AUG, which speci es methionine, the rst amino acid used in the translation process. (Occasionally, GUG is recognized as an initiation codon.) initiation complex The complex formed for initiation of translation. It consists of the 30S ribosomal subunit, messenger RNA, N-formyl-methionine transfer RNA, and three initiation factors. initiation factors (IFI, IF2, IF3) Proteins (prokaryotic with eukaryotic analogues) required for the proper initiation of translation. initiator element (inr) A CT-rich area found in RNA polymerase II promoters without TATA boxes. initiator proteins Proteins that recognize the origin of replication on a replicon and take part in primosome construction. insertion mutagenesis Change in gene action due to an insertion event that either changes a gene directly or disrupts control mechanisms. insertion sequences (IS) Small, simple transposons. See transposable genetic element. inside marker The middle locus of three linked loci. intercalary heterochromatin Heterochromatin, other than centromeric heterochromatin, dispersed throughout eukaryotic chromosomes. intergenic suppression A mutation at a second locus that apparently restores the wild-type phenotype to a mutant at a rst locus. interkinesis The abbreviated interphase that occurs between meiosis I and II. No DNA replication occurs here. internal ribosome entry site Sequence in eukaryotic messenger RNAs that allows ribosomes to initiate translation at a point other than the 5 cap. interphase The metabolically active, nondividing stage of the cell cycle.
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interpolar microtubules Microtubules extending from one pole of the spindle and overlapping spindle bers from the other pole, but not in contact with kinetochores. interrupted mating A mapping technique in which bacterial conjugation is disrupted after speci ed time intervals. intersex An organism with external sexual characteristics of both sexes. intervening sequences (introns) DNA sequences within a gene that are transcribed but removed prior to translation. intra-allelic complementation The restoration of activity in an enzyme made of subunits in a heterozygote of two mutants that, when homozygous, are not active. Caused by the interaction of the subunits in the protein. intragenic suppression A second change within a mutant gene that results in an apparent restoration of the original phenotype. intron See intervening sequences. inversion The replacement of a section of a chromosome in the reverse orientation. inverted repeat sequence A nucleotide sequence read in opposite orientations on the same double helix. in vitro Biological or chemical work done in the test tube (literally, in glass ) rather than in living systems. IS elements See insertion sequences. isochromosome A chromosome with two genetically and morphologically identical arms. isozymes Different electrophoretic forms of the same enzyme. Unlike allozymes, isozymes are due to differing subunit con gurations rather than allelic differences. junctional diversity Variability in immunoglobulins caused by variation in the exact crossover point during V-J,V-D, and D-J joining. kappa particles The bacterialike particles that give a Paramecium the killer phenotype. karyokinesis The process of nuclear division. See cytokinesis. karyotype The chromosome complement of a cell. See idiogram. kinesin Microtubule motor protein. kinetochore The chromosomal attachment point for the spindle bers, located on the centromere. kinetochore microtubules Microtubules radiating from the centrosome and attached to kinetochores of chromosomes during mitosis and meiosis. kin selection The mode of natural selection that acts on an individual s inclusive tness.
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Klenow fragment Proteolytic fragment obtained by treatment with a protease, or protein-cleaving enzyme of E. coli DNA polymerase I with both 5 3 polymerase activity and 3 5 exonuclease activity. (It has been studied extensively because it has been easier for X-ray crystallographers and biochemists to work with this fragment than with the whole enzyme.) knockout mice Transgenic mice that have been made homozygous for a nonfunctioning allele at a particular locus. lac operon The inducible operon, including three loci involved in the uptake and breakdown of lactose. ladder gel See stepladder gel. lagging strand Strand of DNA being replicated discontinuously. lampbrush chromosomes Chromosomes of amphibian oocytes having loops that are suggestive of a lampbrush. leader The length of messenger RNA from the 5 end to the initiation codon, AUG. leader peptide gene A small gene within the attenuator control region of a repressible amino acid operon. Translation of the gene tests the concentration of amino acids in the cell. leader transcript The messenger RNA transcribed by the attenuator region of a repressible amino acid operon. The transcript is capable of several alternative stem-loop structures, depending on the translation of a short leader peptide gene. leading strand Strand of DNA being replicated continuously. leptonema (leptotene stage) The rst stage of prophase I of meiosis, in which chromosomes become distinct. lethal-equivalent alleles Alleles whose summed effect is that of lethality for example, four alleles, each of which would be lethal 25% of the time (or to 25% of their bearers), are equivalent to one lethal allele. leucine zipper Con guration of a DNAbinding protein in which leucine residues on two helices interdigitate, in zipper fashion, to stabilize the protein. leukemia Cancer of the bone marrow resulting in excess production of leukocytes. level of signi cance The probability value in statistics used to reject the null hypothesis. linkage The association of loci on the same chromosome. linkage disequilibrium The condition among alleles at different loci such that allelic combinations in a gamete do not follow the product rule of probability. linkage equilibrium The condition among alleles at different loci such that any allelic combination in a gamete occurs as the product of the frequencies of each allele at its own locus.
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