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linkage groups Associations of loci on the same chromosome. In a species, there are as many linkage groups as there are homologous pairs of chromosomes. linkage number The number of times one strand of a helix coils about the other. linker A small segment of DNA that contains a restriction site. It can be added to bluntended DNA to give that DNA a particular restriction site for cloning. liposomes Transfecting DNA is delivered to target cells by way of these membrane-bound vesicles. locus The position of a gene on a chromosome (plural: loci). lod score method A technique (logarithmic odds) for determining the most likely recombination frequency between two loci from pedigree data. long interspersed elements (LINEs) Sequences of DNA, up to seven thousand base pairs in length, interspersed in eukaryotic chromosomes in many copies. lymphoma Cancer of the lymph nodes and spleen that causes excessive production of lymphocytes. Lyon hypothesis The hypothesis that suggests that the Barr body is an inactivated X chromosome. lysate The contents released from a lysed cell. lysis The breaking open of a cell by the destruction of its wall or membrane. lysogenic The state of a bacterial cell that has an integrated phage (prophage) in its chromosome. major histocompatibility complex A group of highly polymorphic genes whose products appear on the surfaces of cells, imparting to them the property of self (belonging to that organism). Some other functions are also involved. mapping The process of locating the positions of genes on chromosomes. mapping function The mathematical relationship between measured map distance in a given experiment and the actual recombination frequency. map unit The distance equal to 1% recombination between two loci. mate-killer A Paramecium phenotype induced by intracellular bacterialike mu particles. maternal effect The effect of the maternal parent s genotype on the phenotype of her offspring. maternal-effect gene A gene expressed in maternal tissue that in uences a developing embryo. mating type In many species of microorganisms, individuals can be divided into two mating types. Mating can take place only between individuals of opposite mating types due to the interaction of cell surface components.
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maturation-promoting factor (MPF) A protein complex of cyclin B and p34cdc2 that initiates mitosis during the cell cycle. Also called the mitosis-promoting factor. mean The arithmetic average; the sum of the data divided by the sample size. mean tness of the population, W The sum of the tnesses of the genotypes of a population weighted by their proportions; hence, a weighted mean tness. meiosis The nuclear process in diploid eukaryotes that results in gametes or spores with only one member of each original homologous pair of chromosomes per nucleus. meiotic drive See gametic selection. merozygote A bacterial cell having a second copy of a particular chromosomal region in the form of an exogenote. messenger RNA (mRNA) A complementary copy of a gene that is translated into a polypeptide at the ribosome. metacentric chromosome A chromosome whose centromere is located in the middle. metafemale A fruit y with an X/A ratio greater than unity. metagon An RNA necessary for the maintenance of mu particles in Paramecium. metamale A fruit y with X/A ratio below 0.5. metaphase The stage of mitosis or meiosis in which spindle bers are attached to kinetochores, and the chromosomes are positioned in the equatorial plane of the cell. metaphase plate The plane of the equator of the spindle into which chromosomes are positioned during metaphase. metastasis The migration of cancerous cells to other parts of the body. metrical variation See continuous variation. microsatellite DNA Repeats of very short sequences of DNA, such as CACACACA, dispersed throughout the eukaryotic genome. The loci can be studied by polymerase chain reaction ampli cation. microtubule organizing center Active center from which microtubules are organized. The spindle is organized by the centrosome, which may or may not contain a centriole. microtubules Hollow cylinders made of the protein tubulin ( and subunits) that make up, among other things, the spindle bers. mimicry A phenomenon in which an individual gains an advantage by looking like the individuals of a different species. minimal medium A culture medium for microorganisms that contains the minimal necessities for growth of the wild-type. mismatch repair A form of excision repair initiated at the sites of mismatched bases in DNA. missense mutation Mutations that change a codon for one amino acid into a codon for a different amino acid.
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mitochondrion The eukaryotic cellular organelle in which the Krebs cycle and electron transport reactions take place. mitosis The nuclear division producing two daughter nuclei identical to the original nucleus. mitosis-promoting factor See maturationpromoting factor (MPF). mitotic apparatus See spindle. mixed families Groups of four codons sharing their rst two bases and coding for more than one amino acid. modern linkage map A chromosomal map based on the positions of RFLP markers along its length. molecular chaperone A protein that aids in the folding of a second protein. The chaperone prevents proteins from forming structures that would be inactive. molecular evolutionary clock A measurement of evolutionary time in nucleotide substitutions per year. molecular imprinting The phenomenon in which there is differential expression of a gene depending on whether it was maternally or paternally inherited. molecular mimicry The situation in which one type of molecule resembles another type in order to function. For example, the prokaryotic ribosomal release factors, RF1 and RF2, mimic the structure of a transfer RNA. monocistronic Usually referring to a messenger RNA that carries the information for only one gene (cistron). monoclonal antibody The antibody from a clone of cells producing the same antibody. An individual with multiple myeloma usually produces monoclonal antibodies. monohybrids Offspring of parents that differ in only one genetic characteristic. Usually implies heterozygosity at a single locus under study. monosomic A diploid cell missing a single chromosome. monovalent A single chromosome composed of two sister chromatids. Equivalent to a dyad. morphogen A substance transported into or produced in a developing embryo that diffuses to form a gradient that helps determine cell differentiation. morphological species concept The idea that organisms are classi ed in the same species if they appear similar. mosaicism The condition of being a mosaic. See mosaics. mosaics Individuals made up of two or more cell lines in which the cells originated in the same zygote. mRNA See messenger RNA (mRNA). M llerian mimicry A form of mimicry in which noxious species evolve to resemble each other. multihybrid An organism heterozygous at numerous loci.
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