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multinomial expansion The terms generated when a multinomial is raised to a power. mu particles Bacterialike particles found in the cytoplasm of Paramecium that impart the mate-killer phenotype. mutability The ability to change. mutants Alternative phenotypes to the wild-type; the phenotypes produced by alternative alleles. mutation The process by which a gene or chromosome changes structurally; the end result of this process. mutation rate The proportion of mutations per cell division in bacteria or single-celled organisms or the proportion of mutations per gamete in higher organisms. mutator mutations Mutations of DNA polymerase that increase the overall mutation rate of a cell or of an organism. muton A term Benzer coined for the smallest mutable site within a cistron. natural selection The process in nature whereby one genotype leaves more offspring than another genotype because of superior life history attributes such as survival or fecundity. negative interference The phenomenon whereby a crossover in a particular region facilitates the occurrence of other apparent crossovers in the same region of the chromosome. N-end rule The life span of a protein is determined by its amino-terminal (N-terminal) amino acid. neo-Darwinism The merger of classical Darwinian evolution with population genetics. neoplasm New growth of abnormal tissue. neutral gene hypothesis The hypothesis that most genetic variation in natural populations is not maintained by natural selection. NF See fundamental number. nickase See DNA gyrase. noncoding strand See anticoding strand. nondisjunction The failure of a pair of homologous chromosomes to separate properly during meiosis. nonhistone proteins The proteins remaining in chromatin after the histones are removed. nonparental ditype (NPD) A spore arrangement in Ascomycete fungi that contains only the two recombinant-type ascospores (assuming two segregating loci). nonparentals See recombinants. nonrecombinants In mapping studies, offspring that have alleles arranged as in the original parents. nonsense codon One of the messenger RNA sequences (UAA, UAG, UGA) that signals the termination of translation. nonsense mutations Mutations that change a codon for an amino acid into a nonsense codon.
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normal distribution Any of a family of bell-shaped frequency curves whose relative positions and shapes are de ned on the basis of the mean and standard deviation. northern blotting A gel transfer technique used for RNA. See Southern blotting. N segments Sequences of nucleotides added in a template-free fashion at the joining junctions of heavy-chain antibody genes. nuclease-hypersensitive site A region of a eukaryotic chromosome that is speci cally vulnerable to nuclease attack because it is not wrapped as nucleosomes. nucleolar organizer The chromosomal region around which the nucleolus forms; site of tandem repeats of the major ribosomal RNA gene. nucleolus The globular, nuclear organelle formed at the nucleolar organizer. Site of ribosome construction. nucleoprotein The substance of eukaryotic chromosomes consisting of proteins and nucleic acids. nucleoside A sugar-base compound that is a nucleotide precursor. Nucleotides are nucleoside phosphates. nucleosomes Arrangements of DNA and histones forming regular spherical structures in eukaryotic chromatin. nucleotide Subunits that polymerize into nucleic acids (DNA or RNA). Each nucleotide consists of a nitrogenous base, a sugar, and one or more phosphate groups. nucleotide excision repair The DNA excision repair mechanism responsible for repairing thymine dimers and other lesions. Enzymes excise a short segment of one of the DNA strands and then repair and ligate the DNA. null hypothesis The statistical hypothesis that there are no differences between observed data and those data expected based on the assumption of no experimental effect. nullisomic A diploid cell missing both copies of the same chromosome. numerator elements Genes on the X chromosome in Drosophila that regulate the sex switch gene (sxl) to the on condition (femaleness). Refers to the numerator of the X/A genic balance equation. nutritional-requirement mutants See auxotrophs. Okazaki fragments Segments of newly replicated DNA produced during discontinuous DNA replication. oncogene Genes capable of transforming a cell. They are found in the active state in retroviruses and transformed cells and in the inactive state in nontransformed cells, in which they are called proto-oncogenes. one-gene-one-enzyme hypothesis Hypothesis of Beadle and Tatum that states that one gene controls the production of one enzyme. Later modi ed to the concept that one cistron controls the production of one polypeptide.
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oogenesis The process of ovum formation in female animals. oogonia Cells in females that produce primary oocytes by mitosis. open reading frames (ORFs) Sequence of codons between the initiation and termination codons in a gene. operator A DNA sequence recognized by a repressor protein or repressor-corepressor complex. When the operator is complexed with the repressor, transcription is prevented. operon A sequence of adjacent genes all under the transcriptional control of the same operator. origin recognition complex (ORC) A complex of six proteins that bind to the eukaryotic autonomously replicating sequences (ARS). Needed for the initiation of DNA replication in concert with other proteins. outbreeding The mating of genetically unrelated individuals. ovum Egg. The one functional product of each meiosis in female animals. pachynema (pachytene stage) The stage of prophase I of meiosis in which chromatids are rst distinctly visible. palindrome A sequence of words, phrases, or nucleotides that reads the same regardless of the direction from which one starts; the sites of recognition of type II restriction endonucleases. panmictic Referring to unstructured (random mating) populations. paracentric inversion A chromosomal inversion that does not include the centromere. paramecin A toxin liberated by a killer Paramecium. parameters Measurements of attributes of a population; denoted by Greek letters. parapatric speciation Speciation in which reproductive isolating mechanisms evolve when a population enters a new niche or habitat within the range of the parent species. parasegment The rst series of segments that form in a developing insect embryo; they form after about 5.5 hours in the developing Drosophila embryo. parental ditype (PD) A spore arrangement in Ascomycete fungi that contains only the two nonrecombinant-type ascospores. parental imprinting See molecular imprinting. parentals See nonrecombinants. parthenogenesis The development of an individual from an unfertilized egg that did not arise by meiotic chromosomal reduction. partial digest A restriction digest that has not been allowed to go to completion, and thus contains pieces of DNA that have restriction endonuclease sites that have not been cleaved.
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