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product rule The rule that states that the probability that two independent events will both occur is the product of their separate probabilities. progeny testing Breeding of offspring to determine their genotypes and those of their parents. prokaryotes Organisms that lack true nuclei. promoter A DNA region that RNA polymerase binds to in order to initiate transcription. proofread Technically, to read for the purpose of detecting errors for later correction. DNA polymerase has 3 to 5 exonuclease activity, which it uses during polymerization to remove incorrect nucleotides it has recently added. prophage A temperate phage replicating with the host that can later initiate the lytic cycle. prophase The initial stage of mitosis or meiosis in which chromosomes become visible and the spindle apparatus forms. proplastid Mutant plastids that do not grow and develop into chloroplasts. propositus (proposita) The person through whom a pedigree was discovered. proteasome A barrel-shaped cellular organelle for protein breakdown involving the ubuiquitin pathway. proteome From proteins of the genome; the complete set of proteins from a particular genome. It is the protein analogue to genome. proteomics The study of the complete set of proteins from a particular genome. It is the protein analogue to genomics. proto-oncogene A cellular oncogene in an untransformed cell. prototrophs Strains of organisms that can survive on minimal medium. pseudoalleles Genes that are functionally but not structurally allelic. Within gene families, pseudoalleles are alleles that are not expressed. pseudoautosomal gene A gene that occurs on both sex-determining heteromorphic chromosomes. pseudodominance The phenomenon in which a recessive allele shows itself in the phenotype when only one copy of the allele is present, as in hemizygous alleles or in deletion heterozygotes. P (peptidyl) site The site on the ribosome occupied by the peptidyl-tRNA just before peptide bond formation. punctuated equilibrium The evolutionary process involving long periods without change (stasis) punctuated by short periods of rapid speciation. Punnett square A diagrammatic representation of a particular cross used to predict the progeny of the cross. purines Nitrogenous bases of which guanine and adenine are found in DNA and RNA.
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pyrimidines Nitrogenous bases of which thymine is found in DNA, uracil in RNA, and cytosine in both. quantitative inheritance The mechanism of genetic control of traits showing continuous variation. quantitative trait loci Chromosomal regions contributing to the inheritance of a quantitative trait. These regions may contain one or more polygenes that contribute to the phenotype. quantitative variation See continuous variation. quaternary structure The association of polypeptide subunits to form the nal structure of a protein. random genetic drift Changes in allelic frequency due to sampling error. random mating The mating of individuals in a population such that the union of individuals with the trait under study occurs according to the product rule of probability. random strand analysis Mapping studies in organisms that do not keep all the products of meiosis together. read-through Transcription or translation beyond the normal termination signals in DNA or RNA, respectively. realized heritability Heritability measured by a response to selection. recessive An allele (or phenotype) that does not express itself in the heterozygous condition. reciprocal cross A cross with the phenotype of each sex reversed as compared with the original cross. Made to test the role of parental sex on inheritance pattern. reciprocal translocation A chromosomal con guration in which the ends of two nonhomologous chromosomes are broken off and become attached to the nonhomologues. recombinant DNA technology Techniques of gene cloning. Recombinant DNA refers to the hybrid of foreign and vector DNA. See gene cloning. recombinant plasmid A plasmid that contains an inserted piece of foreign DNA. recombinants In mapping studies, offspring with allelic arrangements made up of a combination of the original parental alleles. recombination The nonparental arrangement of alleles in progeny that can result from either independent assortment or crossing over. recombination nodule Proteinaceous nodules found on bivalents during zygonema and pachynema associated with crossing over. recon A term Benzer coined for the smallest recombinable unit within a cistron. reductional division The rst meiotic division. It reduces the number of chromosomes and centromeres to half that in the original cell.
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regional centromere The type of centromere found in higher eukaryotes that can accommodate several spindle microtubules. regulator gene A gene primarily involved in control of the production of another gene s product. reinitiation The initiation of translation by a ribosome that has just completed translation of a region of the messenger RNA upstream of the current point of initiation. relative Darwinian tness See tness. relaxed mutant A mutant that does not exhibit the stringent response under amino acid starvation. release factors (RFl and RF2) Proteins in prokaryotes responsible for termination of translation and release of the newly synthesized polypeptide when a nonsense codon appears in the A site of the ribosome. Replaced by eRF in eukaryotes. repetitive DNA DNA made up of copies of the same nucleotide sequence. replica-plating A technique to rapidly transfer microorganism colonies to numerous petri plates. replication-coupling assembly factor A protein complex in fruit ies that assembles new nucleosomes. replicons A replicating genetic unit including a length of DNA and its site for the initiation of replication. replisome The DNA-replicating structure at the Y-junction, consisting of two DNA polymerase III enzymes and a primosome (primase and DNA helicase). reporter systems Genetic constructs that allow an investigator to determine that a speci c locus is active by measuring the phenotypic expression of an associated locus, such as the luciferase reporter, which glows if watered with luciferin. repressible system A coordinated group of enzymes involved in a synthetic pathway (anabolic) is repressible if excess quantities of the end product of the pathway lead to the termination of transcription of the genes for the enzymes. These systems are primarily prokaryotic operons. repressor The protein product of a regulator gene that acts to control transcription of inducible and repressible operons. reproductive isolating mechanisms Environmental, behavioral, mechanical, and physiological barriers that prevent two individuals of different populations from producing viable progeny. reproductive success The relative production of offspring by a particular genotype. repulsion Allelic arrangement in which each homologous chromosome has mutant and wild-type alleles. resistance transfer factor Infectious transfer part of R plasmids.
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