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structural alleles Alleles whose alterations include some of the same base pairs. submetacentric chromosome A chromosome whose centromere lies between its middle and its end, but closer to the middle. subtelocentric chromosome A chromosome whose centromere lies between its middle and its end, but closer to the end. sum rule The rule that states that the probability that two mutually exclusive events will occur is the sum of the probabilities of the individual events. supercoiling Negative or positive coiling of double-stranded DNA that differs from the relaxed state. supergenes Several loci, which usually control related aspects of the phenotype, in close physical association. suppressor gene A gene that, when mutated, apparently restores the wild-type phenotype to a mutant of another locus. surveillance mechanism Used to describe mechanisms that oversee checkpoints in the cell cycle, points in which the process can be halted if certain conditions are not met. survival of the ttest In evolutionary theory, survival of only those organisms best able to obtain and utilize resources (the fittest). This phenomenon is the cornerstone of Darwin s theory of evolution. Svedberg unit A unit of sedimentation during centrifugation. Abbreviation is S, as in 50S. swivelase See DNA gyrase. sympatric speciation Speciation in which reproductive isolating mechanisms evolve within the range and habitat of the parent species. This type of speciation may be common in parasites. synapsis The point-by-point pairing of homologous chromosomes during zygonema or in certain dipteran tissues that undergo endomitosis. synaptonemal complex A proteinaceous complex that apparently mediates synapsis during the zygotene stage and then disintegrates. syncitium A cell that has many nuclei not separated by cell membranes. synexpression group A group of eukaryotic genes that are involved in the same function or pathway and are induced together. synteny test A test that determines whether two loci belong to the same linkage group by observing concordance in hybrid cell lines. synthetic medium A chemically de ned substrate that microorganisms are grown upon. TACTAAC box A consensus sequence surrounding the lariat branch point of eukaryotic messenger RNA introns.
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TATA-binding protein A protein, part of TFIID, that binds the TATA consensus sequence in eukaryotic promoters. TATA box An invariant DNA sequence at about 25 in the promoter region of eukaryotic genes; analogous to the Pribnow box in prokaryotes. tautomeric shift Reversible shifts of proton positions in a molecule. The bases in nucleic acids shift between the keto and enol forms or between the amino and imino forms. TBP-associated factors (TAFs) Proteins that bind with the TATA-binding protein to form TFIID. They aid in the selectivity of TFIID. T-cell receptors Surface proteins of T cells that allow the T cells to recognize host cells that have been infected. telocentric chromosome A chromosome whose centromere lies at one of its ends. telomerase An enzyme that adds telomeric sequences to the ends of eukaryotic chromosomes. telomeres The ends of linear chromosomes. telophase The terminal stage of mitosis or meiosis in which chromosomes uncoil, the spindle breaks down, and cytokinesis usually occurs. telson The posterior end of the arthropod embryo, where the end of the alimentary canal is located. temperate phage A phage that can enter into lysogeny with its host. temperature-sensitive mutant An organism with an allele that is normal at a permissive temperature but mutant at a restrictive temperature. template A pattern serving as a mechanical guide. In DNA replication, each strand of the duplex acts as a template for the synthesis of a new double helix. template strand See anticoding strand. terminator sequence A sequence in DNA that signals the termination of transcription to RNA polymerase. terminator stem A con guration of the leader transcript that signals transcriptional termination in attenuator-controlled amino acid operons. tertiary structure The further folding of a protein, bringing helices and sheets into three-dimensional arrangements. testcross The cross of an organism with a homozygous recessive organism. testing of hypotheses The determination of whether to reject or fail to reject a proposed hypothesis based on the likelihood of the experimental results. testis-determining factor (TDF) General term for the gene determining maleness in human beings (Tdf in mice). tetrads The meiotic con guration of four chromatids rst seen in pachynema. There is one tetrad (bivalent) per homologous pair of chromosomes.
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tetranucleotide hypothesis The hypothesis, based on incorrect information, that DNA could not be the genetic material because its structure was too simple that is, that repeating subunits contain one copy each of the four DNA nucleotides. tetraploids Organisms with four whole sets of chromosomes. tetratype (TT) A spore arrangement in Ascomycete fungi that consists of the two parental and two recombinant spores. theta structure An intermediate structure formed during the replication of a circular DNA molecule. three-point cross A cross involving three loci. thymine See pyrimidines. t-loop A loop that forms at the end of mammalian telomeres by the interdigitation of the 3 free end into the DNA double helix. topoisomerase An enzyme that can relieve (or create) supercoiling in DNA by creating transitory breaks in one (type I) or both (type II) strands of the helical backbone. topoisomers Forms of DNA with the same sequence but differing in their linkage number (coiling). totipotent The state of a cell that can give rise to any and all adult cell types, as compared with a differentiated cell whose fate is determined. trailer The length of messenger RNA from the nonsense codon to the 3 end or, in polycistronic messenger RNAs, from a nonsense codon to the next gene s leader. trans Meaning across and referring usually to the geometric con guration of mutant alleles across from each other on a homologous pair of chromosomes. trans-acting Referring to mutations of, for example, a repressor gene, that act through a diffusible protein product; the normal mode of action of most recessive mutations. transcription The process whereby RNA is synthesized from a DNA template. transcription factors Eukaryotic proteins that aid RNA polymerase in recognizing promoters. See general transcription factors and speci c transcription factors. transducing particle A defective phage, carrying part of the host s genome. transduction A process whereby a cell can gain access to and incorporate foreign DNA brought in by a viral particle. transfection The introduction of foreign DNA into eukaryotic cells. transfer operon (tra) Sequence of loci that impart the male (F-pili-producing) phenotype on a bacterium. The male cell can transfer the F plasmid to an F cell. transfer RNA (tRNA) Small RNA molecules that carry amino acids to the ribosome for polymerization.
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