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transformation A process whereby prokaryotes take up DNA from the environment and incorporate it into their genomes, or the conversion of a eukaryotic cell into a cancerous one. transgenic Eukaryotic organisms that have taken up foreign DNA. transition mutation A mutation in which a purine-pyrimidine base pair replaces a base pair in the same purine-pyrimidine relationship. translation The process of protein synthesis wherein the nucleotide sequence in messenger RNA determines the primary structure of the protein. translocase (EF-G) Elongation factor in prokaryotes necessary for proper translocation at the ribosome during the translation process. Replaced by eEF2 in eukaryotes. translocation A chromosomal con guration in which part of a chromosome becomes attached to a different chromosome. Also a part of the translation process in which the messenger RNA is shifted one codon in relation to the ribosome. translocation channel (translocon) A protein-lined pore or channel in a membrane through which nascent proteins are transported during translation. transposable genetic element A region of the genome, anked by inverted repeats, a copy of which can be inserted at another place; also called a transposon or a jumping gene. transposon See transposable genetic element. transversion mutation A mutation in which a purine replaces a pyrimidine, or vice versa. trihybrid An organism heterozygous at three loci. triploids Organisms with three whole sets of chromosomes. trisomic A diploid cell with an extra chromosome. tRNA See transfer RNA (tRNA). trp RNA-binding attenuation protein (TRAP) Protein that can bind to the attenuation region of the messenger RNA of the tryptophan operon in Bacillus subtilis, causing a terminator stem to form and halting further transcription. true heritability See heritability. tumor Abnormal growth of tissue. tumor-suppressor genes Genes that normally prevent unlimited cellular growth. When both copies of the gene are mutated, cellular transformation follows. Examples are the p53 gene and the genes for retinoblastoma and Wilm s tumor. two-point cross A cross involving two loci.
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type I error In statistics, the rejection of a true hypothesis. type II error In statistics, the accepting of a false hypothesis. typological thinking The concept that organisms of a species conform to a speci c norm. In this view, variation is considered abnormal. ubiquitin A peptide of twenty-six amino acid residues that enzymes attach to proteins that the proteasome will degrade. unequal crossing over Nonreciprocal crossing over caused by the mismatching of homologous chromosomes. Usually occurs in regions of tandem repeats. uninemic chromosome A chromosome consisting of one DNA double helix. unique DNA A length of DNA with no repetitive nucleotide sequences. unmixed families Groups of four codons sharing their rst two bases and coding for the same amino acid. unusual bases Other bases, in addition to adenine, cytosine, guanine, and uracil, found primarily in transfer RNAs. UP element See upstream element. upstream A convention on DNA related to the position and direction of transcription by RNA polymerase (5 3 ). Downstream (or 3 to) is in the direction of transcription whereas upstream (5 to) is in the direction from which the polymerase has come. upstream element A sequence of about twenty AT-rich bases centered at 50 in promoters of prokaryotic genes that are expressed strongly. uracil See pyrimidines. variable-number-of-tandem-repeats (VNTR) loci Loci that are hypervariable because of tandem repeats. Presumably, variability is generated by unequal crossing over. variance The average squared deviation about the mean of a set of data. variegation Patchiness; a type of position effect that results when particular loci are contiguous with heterochromatin. virion A virus particle. viroids Bare RNA particles that are plant pathogens. V(D)J joining The process of joining variable, diversity, and joining gene segments (V-J,V-D, or D-J joining) in the formation of a functioning immunoglobulin gene. Wahlund effect The effect of subdivision on a population, causing it to contain fewer heterozygotes than predicted despite the fact that all subdivisions are in Hardy-Weinberg proportions. western blotting A technique for probing for a particular protein using antibodies. See Southern blotting.
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whole-genome shotgun method A method of sequencing entire genomes by breaking up the genomes into small pieces, sequencing the pieces, and then using computers to establish order by overlapping the sequences. wild-type The phenotype of a particular organism when it is rst seen in nature. Wilm s tumor A childhood kidney cancer caused by the inactivation of an anti-oncogene. wobble Referring to the reduced constraint over the third base of an anticodon as compared with the other bases, thus allowing additional complementary base pairings. xeroderma pigmentosum A disease in human beings caused by a defect in the UV mutation repair system. X-inactivation center (XIC) Locus at which inactivation is initiated on the X chromosome in mammals. X linked See sex-linked. X-ray crystallography A technique, using X rays, to determine the atomic structure of molecules that have been crystallized. yeast arti cial chromosome (YAC) Originating from a bacterial plasmid, a YAC contains additionally a yeast centromeric region (CEN) and a yeast origin of DNA replication (ARS). YACs are capable of including large pieces of foreign DNA during cloning. Y-junction The point of active DNA replication where the double helix opens up so that each strand can serve as a template. Y linked Inheritance pattern of loci located on the Y chromosome only. Also refers to the loci themselves. Z DNA A left-handed form of DNA found under physiological conditions in short GC segments that are methylated. It may be involved in regulating gene expression in eukaryotes. ZFY gene Originally believed to be the human male sex-switch gene, located on the short arm of the Y chromosome. ZFY stands for zinc finger on the Y chromosome. zinc nger Con guration of a DNA-binding protein that resembles a nger with a base, usually cysteines and histidines, binding a zinc ion. Discovered in a transcription factor in Xenopus. zygonema (zygotene stage) The stage of prophase I of meiosis in which synapsis occurs. zygotic induction The beginning of vegetative growth when a prophage is passed into an F cell during conjugation. zygotic selection The forces acting to cause differential mortality of an organism at any stage (other than gametes) in its life cycle.
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