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Tamarin: Principles of Genetics, Seventh Edition
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II. Mendelism and the Chromosomal Theory
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6. Linkage and Mapping in Eukaryotes
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n 10 December each year, the king of Sweden awards the Nobel Prizes at the Stockholm Concert Hall. The date is the anniversary of Alfred Nobel s death. Awards are given annually in physics, chemistry, medicine and physiology, literature, economics, and peace. In 2000, each award was worth $900,000, although an award sometimes goes to two or three recipients. The prestige is priceless. Winners of the Nobel Prize are chosen according to the will of Alfred Nobel, a wealthy Swedish inventor and industrialist, who held over three hundred patents when he died in 1896 at the age of sixty-three. Nobel developed a detonator and processes for detonation of nitroglycerine, a substance invented by Italian chemist Ascanio Sobrero in 1847. In the form Nobel developed, the explosive was patented as dynamite. Nobel also invented several other forms of explosives. He was a benefactor of Sobrero, hiring him as a consultant and paying his wife a pension after Sobrero died. Nobel believed that dynamite would be so destructive that it would serve as a deterrent to war. Later, realizing that this would not come to pass, he instructed that his fortune be invested and the interest used to fund the awards. The rst prizes were awarded in 1901. Each award consists of a diploma, medal, and check.
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American, British, German, French, and Swedish citizens have earned the most prizes (table 1). Table 2 features some highlights of Nobel laureate achievements in genetics.
The Nobel medal. The medal is half a pound of 23-karat gold, measures about 2 1/2 inches across, and has Nobel s face and the dates of his birth and death on the front. The diplomas that accompany the awards are individually designed.
(Reproduced by permission of the Nobel Foundation.)
Table 1 Distribution of Nobel Awards to the Top Five Recipient Nations (Including 2000 Winners)
Physics Chemistry Medicine and Physiology Peace Literature Economics Total
United States Britain Germany France Sweden
77 20 18 12 4
46 24 27 7 4
88 25 15 7 7
20 9 4 8 5
9 8 6 12 7
30 5 1 1 2
270 91 71 47 29
Tamarin: Principles of Genetics, Seventh Edition
II. Mendelism and the Chromosomal Theory
6. Linkage and Mapping in Eukaryotes
The McGraw Hill Companies, 2001
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Table 2 Some Nobel Laureates in Genetics (Medicine and Physiology; Chemistry)
Name Year Nationality Cited for
Thomas Hunt Morgan Hermann J. Muller George W. Beadle Edward L. Tatum Joshua Lederberg Severo Ochoa Arthur Kornberg Francis H. C. Crick James D. Watson Maurice Wilkins Fran ois Jacob Andr Lwoff Jacques Monod Peyton Rous Robert W. Holley H. Gobind Khorana Marshall W. Nirenberg Max Delbr ck Alfred Hershey Salvador Luria Renato Dulbecco Howard Temin David Baltimore Werner Arber Hamilton Smith Daniel Nathans Walter Gilbert Frederick Sanger Paul Berg Baruj Benacerraf Jean Dausset George Snell Aaron Klug Barbara McClintock Cesar Milstein Georges Koehler Niels K. Jerne Susumu Tonegawa J. Michael Bishop Harold E.Varmus Thomas R. Cech Sidney Altman Kary Mullis Michael Smith Richard Roberts Phillip Sharp E. B. Lewis
1933 1946 1958 1958 1958 1959 1959 1962 1962 1962 1965 1965 1965 1966 1968 1968 1968 1969 1969 1969 1975 1975 1975 1978 1978 1978 1980 1980 1980 1980 1980 1980 1982 1983 1984 1984 1984 1987 1989 1989 1989 1989 1993 1993 1993 1993 1995
USA USA USA USA USA USA USA British USA British French French French USA USA USA USA USA USA USA USA USA USA Swiss USA USA USA British USA USA French USA British USA British/Argentine German British/Danish Japanese USA USA USA Canada USA Canada British USA USA
Discovery of how chromosomes govern heredity X-ray inducement of mutations Genetic regulation of biosynthetic pathways Bacterial genetics Discovery of enzymes that synthesize nucleic acids Discovery of the structure of DNA
Regulation of enzyme biosynthesis
Tumor viruses Unraveling of the genetic code
Viral genetics
Tumor viruses Discovery of reverse transcriptase Discovery and use of restriction endonucleases
Techniques of sequencing DNA Pioneer work in recombinant DNA Genetics of immune reactions
Crystallographic work on protein-nucleic acid complexes Transposable genetic elements Immunogenetics
Antibody diversity Proto-oncogenes Enzymatic properties of RNA Polymerase chain reaction Site-directed mutagenesis Discovery of intervening sequences in RNA Genes control development
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