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II. Mendelism and the Chromosomal Theory
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6. Linkage and Mapping in Eukaryotes
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16. A certain haploid strain of yeast was de cient for the synthesis of the amino acids tryptophan (try ) and methionine (met ). It was crossed to the wild-type, and meiosis occurred. One dozen asci were analyzed for their tryptophan and methionine requirements. The following results, with the inevitable lost spores, were obtained:
1 2 3 4 5 6 7 8 9 10 11 12 try try try try try try try try try try met met met met met met met met met met try try try try try try try try try try met met met met met met met met met met try try try try try met met try try try try try try try try try try try met met met met met met met met met met met
1 2 3 4 5 6 7 8 9 10 11 12
f f f f f f f f f f
f f f f f f f f f
f f f f f f f
f f f f
f f f f f f
f f f f f f f f f
f f f f f f f f f
f f f f f f f f f
a. Classify each ascus as to segregational type, and note which asci cannot be classi ed. b. Map the chromosome containing the f locus with all the relevant measurements. 19. Draw ten of the remaining twenty-eight ascus patterns not included in table 6.6.To which of the seven major categories of table 6.7 does each belong 20. In yeast, the a and b loci are 12 map units apart. Construct a data set to demonstrate this. 21. In Neurospora, the a locus is 12 map units from its centromere. Construct a data set to show this. 22. An ab Neurospora was crossed with an a b form. Meiosis occurred, and 1,000 asci were dissected. Using the classes of table 6.7, the following data resulted:
Class 1 Class 2 Class 3 Class 4 700 0 190 90 Class 5 Class 6 Class 7 5 5 10
met met
a. Classify each ascus as to segregational type (note that some asci may not be classi able). b. Are the genes linked c. If so, how far apart are they 17. In Neurospora, a haploid strain requiring arginine (arg ) is crossed with the wild-type (arg ). Meiosis occurs, and ten asci are dissected with the following results. Map the arg locus.
1 2 3 4 5 6 7 8 9 10 arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg arg
What is the linkage arrangement of these loci 23. Given the following linkage arrangement in Neurospora, construct a data set similar to that in table 6.7 that is consistent with it (cm is centromere). a 15 cm
b 15
24. Determine crossover events that led to each of the seven classes in table 6.7. 25. In Neurospora, a cross is made between ab and a b individuals.The following one hundred ordered tetrads are obtained:
Spores I 1, 2 3, 4 5, 6 7, 8 a b a b ab ab 85 II a b a b ab ab 2 III a b a b ab ab 3 IV a b a b ab ab 2 V a b a b ab ab 3 VI a b ab a b ab 3 VII a b ab ab a b 1 VIII ab a b a b ab 1
18. A haploid strain of Neurospora with fuzzy colony morphology ( f ) was crossed with the wild-type ( f ).Twelve asci were scored.The following results, with the inevitable lost spores were obtained:
a. Are genes a and b linked How do you know b. Calculate the gene-to-centromere distances for a and b.
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