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Tamarin: Principles of Genetics, Seventh Edition
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II. Mendelism and the Chromosomal Theory
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6. Linkage and Mapping in Eukaryotes
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The McGraw Hill Companies, 2001
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Critical Thinking Questions
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35. You have selected three mouse-human hybrid clones and analyzed them for the presence of human chromosomes. You then analyze each clone for the presence or absence of particular human enzymes ( presence of human chromosome or enzyme activity). Based on the following results indicate the probable chromosomal location for each enzyme.
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Human Chromosomes Clone X Y Z 3 7 9 11 15 18 20
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36. Three mouse-human cell lines were scored for the presence ( ) or absence ( ) of human chromosomes, with the results as follows:
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Human Chromosomes Clone A B C 1 2 3 4 5 14 15 18
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If a particular gene is located on chromosome 3, which clones should be positive for the enzyme from that gene
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Enzyme
Clone X Y Z
C R I T I C A L
1. 2.
T H I N K I N G
Q U E S T I O N S
Do three-point crosses in fruit ies capture all the multiple crossovers in a region If 4% of all tetrads have a single crossover between two loci: (a) What is the map distance between these
loci if these are fruit ies (b) What is the proportion of second-division segregants if these are Neurospora (c) What is the proportion of nonparental ditypes if these are yeast
Suggested Readings for chapter 6 are on page B-3.
Tamarin: Principles of Genetics, Seventh Edition
II. Mendelism and the Chromosomal Theory
7. Linkage and Mapping in Prokaryotes and Bacterial Viruses
The McGraw Hill Companies, 2001
LINKAGE AND MAPPING IN PROKARYOTES AND BACTERIAL VIRUSES
STUDY OBJECTIVES
1. To de ne bacteria and bacterial viruses and learn about methods of studying them 149 2. To study life cycles and sexual processes in bacteria and bacteriophages 154, 163 3. To make use of the sexual processes of bacteria and their viruses to map their chromosomes 155, 166
STUDY OUTLINE
Bacteria and Bacterial Viruses in Genetic Research 149 Techniques of Cultivation 150 Bacterial Phenotypes 151 Colony Morphology 151 Nutritional Requirements 151 Resistance and Sensitivity 153 Viral Phenotypes 154 Sexual Processes in Bacteria and Bacteriophages 154 Transformation 154 Conjugation 157 Life Cycles of Bacteriophages 163 Recombination 163 Lysogeny 165 Transduction 165 Specialized Transduction 165 Generalized Transduction 166 Mapping with Transduction 166 Summary 171 Solved Problems 172 Exercises and Problems 172 Critical Thinking Questions 176
Scanning electron micrograph (color enhanced) of an Escherichia coli bacterium with adsorbed T-family bacteriophages (36,000x). ( Oliver Meckes/MPITubingen/Photo Researchers.)
Tamarin: Principles of Genetics, Seventh Edition
II. Mendelism and the Chromosomal Theory
7. Linkage and Mapping in Prokaryotes and Bacterial Viruses
The McGraw Hill Companies, 2001
Bacteria and Bacterial Viruses in Genetic Research
ll organisms and viruses have genes located sequentially in their genetic material; and almost all can undergo recombination between homologous (equivalent) pieces of genetic material. Because recombination can occur, it is possible to map, by analytical methods, the locations and sequence of genes along the chromosomes of all organisms and almost all viruses. In this chapter, the viruses we look at are those that attack bacteria.Through work with bacteria and viruses, we have entered the modern era of molecular genetics, the subject of the next section of this book. Bacteria (including the cyanobacteria, the blue-green algae) are prokaryotes. The prokaryotes also include the archaea, or archaebacteria, a kingdom recognized in 1980. These highly specialized organisms (previously classi ed as bacteria), along with the bacteria and eukaryotes, make up the three domains of life on Earth. The true bacteria can be classi ed according to shape: a spherical bacterium is called a coccus; a rod-shaped bacterium is called a bacillus; and a spiral bacterium is called a spirillum. Prokaryotes do not undergo mitosis or meiosis but simply divide in two after their chromosome (usually only one), most often a circle of DNA, has replicated (see chapter 9). Bacterial viruses do not even divide; they are mass-produced within a host cell.
ested in studying in this chapter the bacterial viruses, bacteriophages, or just phages (Greek: eating) are exclusively genetic material surrounded by a protein coat. Bacteriophages are usually classi ed rst by the type of genetic material (nucleic acid) they have (DNA or RNA, single- or double-stranded), then by structural features of their protein surfaces (capsids) such as type or symmetry and number of discrete protein subunits (capsomeres) in the capsid, and general size. Most bacteriophages are complex, like T2 ( g. 7.1), or made up of a headlike capsule like T2 without the tail appendages, or lamentous. Most contain double-stranded DNA. Bacteriophages are obligate parasites; outside of a host, they are inert molecules. Once their genetic material penetrates a host cell, they can take over the metabolism of that cell and construct multiple copies of themselves. We will discuss details of this and alternative infection
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